Glossary of Key Terms and Concepts

Action Potential (AP)

A temporary change in the electrical membrane potential of a cell, typically a nerve or muscle cell, that leads to an electrical impulse. In the heart, each cardiac action potential triggers a heartbeat by initiating the contraction of the cardiac muscle cells.

Action Potential Duration (APD)

The length of time from the start of a cardiac action potential (depolarization) until repolarization is complete. The APD strongly influences the heart’s refractory period (the period during which cells cannot be re-excited).

Afterdepolarizations

Abnormal voltage fluctuations following or during a cardiac action potential. They can trigger extra beats (arrhythmias) if they reach threshold.

Atrial Fibrillation (AF)

A common cardiac arrhythmia in which the atria (the heart’s upper chambers) quiver or beat irregularly, leading to an irregular and often rapid heartbeat.

ATP-Dependent Na⁺-K⁺ Pump

A membrane protein that uses ATP (energy) to move sodium (Na⁺) and potassium (K⁺) ions across the cell membrane (3 sodium ions out, 2 potassium ions in). Helps maintain the resting membrane potential and ionic gradients essential for heart cell function.

Bradycardia

A slower than normal heart rate (usually fewer than 60 beats per minute in adults).

Calcium (Ca²⁺)-Induced Ca²⁺ Release

In the heart, the influx of calcium through L-type calcium channels triggers a larger release of calcium from the sarcoplasmic reticulum (SR). This rise in intracellular calcium drives cardiac muscle contraction.

Calcium Currents and Channels

Calcium Handling Proteins

Cardiac Arrhythmia

An abnormal heart rhythm. It can be too fast, too slow, or irregular, and arises from disturbances in how impulses are generated or conducted through the heart.

Cardiomyocyte

A heart muscle cell. Cardiomyocytes are specialized for contraction and have unique electrical properties that enable the heartbeat.

Congenital Long QT Syndrome (LQTS)

A hereditary condition in which the QT interval (on the ECG) is prolonged, usually due to mutations in specific ion channel genes. It can cause dangerous arrhythmias.

Conduction Block

Failure of an electrical impulse to propagate normally through the heart. Can be “unidirectional” (the impulse travels in one direction but not the reverse) or “bidirectional” (blocked in both directions). Plays a role in reentry-based arrhythmias.

Coupled Clock Hypothesis

An explanation of pacemaker function in sinoatrial node cells, stating that both membrane currents (the “membrane clock,” such as the funny current I_f) and rhythmic calcium cycling inside the cell (the “calcium clock”) interact to set and regulate the heart’s natural pacing.

Diastole

The phase of the cardiac cycle when the heart muscle relaxes and allows the chambers to fill with blood.

Dispersion of Repolarization

Differences in action potential durations (or repolarization timing) across different regions of the heart muscle. Heightened dispersion can create a substrate for reentrant arrhythmias.

Effective Refractory Period (ERP)

The shortest time interval after an action potential during which a new electrical stimulus cannot trigger another action potential in cardiac cells.

Electrocardiogram (ECG or EKG)

A noninvasive recording of the heart’s electrical activity measured from the body surface. Common features include the P wave (atrial depolarization), QRS complex (ventricular depolarization), and T wave (ventricular repolarization).

Excitation-Contraction Coupling

The sequence of events whereby an action potential in a heart cell leads to calcium influx, which then triggers cellular contraction.

Funny Current (I_f)

A mixed inward current (carried by both sodium and potassium) that activates upon hyperpolarization. Plays a key role in pacemaker activity, particularly in the sinoatrial node.

Gap Junction

Specialized cell-cell connections (made of proteins called connexins) that allow direct electrical communication between adjacent heart cells, enabling coordinated propagation of electrical impulses.

Heart Failure (HF)

A clinical syndrome in which the heart’s ability to pump blood is diminished. HF often involves electrical remodeling, changes in calcium handling, and increased risk of arrhythmias.

Hyperpolarization

A shift of the membrane potential to a more negative value, usually making it harder for a cell to fire an action potential.

Ionic Currents in Cardiac Cells

Late Sodium Current (I_NaLate)

A small but prolonged component of the sodium current that persists beyond the initial fast upstroke. Increased late sodium current can prolong repolarization and predispose cells to arrhythmias.

Myocardial Infarction (MI)

Commonly known as a heart attack. It occurs when blood flow to part of the heart is blocked, causing cell death and often leading to electrical and structural remodeling that may set the stage for arrhythmias.

Na⁺-Ca²⁺ Exchanger (NCX)

A membrane transporter that exchanges 3 sodium ions for 1 calcium ion, moving calcium out of the cell (or in the opposite direction depending on membrane potential and ion gradients). Plays a significant role in shaping the action potential plateau and in generating afterdepolarizations when intracellular calcium is high.

Nernst Equation

A formula used to calculate the equilibrium potential of an ion (e.g., K⁺) across a membrane based on its concentration gradient and charge.

Pacemaker Activity

Automatic (self-initiated) firing of action potentials, typically in sinoatrial node cells, setting the heart rate. Certain disease conditions or ectopic foci can also develop pacemaker-like activity.

Phase 0, 1, 2, 3, 4 of the Cardiac Action Potential

Purkinje Fibers

Specialized conducting fibers in the ventricles that rapidly distribute the electrical impulse to ensure coordinated ventricular contraction. They can also act as subsidiary pacemakers if the primary pacemaker (SA node) fails.

Reentry

An arrhythmia mechanism in which a wave of electrical excitation keeps re-circulating through a region of tissue (e.g., because of conduction block in one direction, slow conduction, and reactivation in the reverse direction). This can sustain rapid tachyarrhythmias.

Repolarization

The return of the cardiac cell membrane potential from a positive (depolarized) value back toward the negative resting level (phase 3 of the action potential).

Restitution (Electrical Restitution)

How action potential duration (APD) adapts following a preceding beat, often studied by looking at APD as a function of the diastolic interval. A steep APD restitution slope can favor arrhythmias (e.g., EADs or alternans).

Sarcoplasmic Reticulum (SR)

A specialized endoplasmic reticulum in muscle cells that stores calcium. Calcium release from the SR initiates contraction, and re-uptake into the SR enables relaxation.

Sinoatrial Node (SAN)

The heart’s primary pacemaker region, located in the right atrium. SAN cells spontaneously depolarize, setting the normal heart rate.

Tachyarrhythmia / Tachycardia

A rapid heart rhythm (often above 100 beats per minute in adults). It can be regular or irregular.

Transmembrane Potential

The voltage difference across a cell’s membrane—inside relative to outside. Heart cell transmembrane potential changes drive the cardiac action potential.

Triggered Activity / Triggered Automaticity

Abnormal impulses occurring when afterdepolarizations (EADs or DADs) reach threshold, initiating extra beats that can degenerate into more dangerous arrhythmias.

Voltage Clamp / Patch Clamp

Laboratory methods for measuring ion currents in isolated cells (or tissue) by controlling or recording the membrane potential. Widely used to identify and quantify specific ion channel behaviors.